Regularidad del escalamiento metabólico en ecosistemas acuáticos

Autores/as

  • L. A. Salcido -Guevara
  • F. Arreguí­n -Sánchez
  • L. Palmeri
  • A. Barausse

DOI:

https://doi.org/10.37543/oceanides.v27i2.113

Palabras clave:

Ecosistema, metabolismo, factor de escalamiento, eficiencia de transferencia

Resumen

Se contrastó la hipótesis de que el metabolismo de un ecosistema sigue una relación de escalamiento análoga a la existente en los organismos. La relación entre el logaritmo de la razón Producción/
Biomasa (B/P) y el nivel trófico (TL) se estimó para 98 modelos tróficos de los ecosistemas acuáticos. Una distribución normal de las pendientes de esta relación produjo un valor modal de 0.64 que es significativamente diferente del valor teórico de 0.75 (p<0.05). Después de realizar una corrección considerando la eficiencia de transferencia entre niveles tróficos, se obtuvo un valor modal de 0.726, el cual fue obtenido a través de un algoritmo de mínimos cuadrados, que generó un valor significativamente (p>0.05) similar al teórico esperado. También se contrastó la hipótesis de existencia de error en ambas variables, logaritmo (B/P) y TL, a través de la técnica de regresión denominada “Reduced Major Axis”, con resultados similares según el valor modal de 0.756, sin diferencia estadísticamente significativa (p>0.05) del valor teórico. Se exploró la existencia de algún
patrón en la distribución geográfica, sin obtenerse relación significativa (p>0.05) con la latitud, o con diferentes regiones del mundo. Las conclusiones son: a) el metabolismo del ecosistema sigue la regla de escalamiento metabólico de 3/4; b) la eficiencia de la transferencia entre TL desempeña un papel relevante, representando los atributos locales del metabolismo del ecosistema; c) no hay una diferencias latitudinal o geográfica. Estos
resultados confirman la existencia de una regularidad en el escalamiento metabólico en ecosistemas acuáticos.

Descargas

Los datos de descargas todavía no están disponibles.

Citas

Allen, K.R. 1971. Relation between production and biomass. J. Fish. Res. Board Can., 28(10): 1573-1581. https://doi.org/10.1139/f71-236

Bohonak, A.J. & K. van der Linde. 2004. RMA: Software for Reduced Major Axis regression, Java version. Available from: http:// www.kimvdlinde.com/professional/rma. html [Accessed 3 February 2012].

Banavar, J.R., A. Maritan & A. Rinaldo.1999. Size and form in efficient transportation networks. Nature, 399: 130-132. https://doi.org/10.1038/20144

Banavar, J.R., A. Maritan & A. Rinaldo.2002. Supply-demand balance and metabolic scaling. PNAS, 99(16): 10506-10509. https://doi.org/10.1073/pnas.162216899

Banavar, J.R., M.E. Moses, J.H. Brown, J. Damuth, A. Rinaldo, R.M. Sibly & A. Maritan. 2010. A general basis for quarter-power scaling in animals. PNAS. 107(36): 15816-15820. https://doi.org/10.1073/pnas.1009974107

Bendoricchio, G. & L. Palmeri. 2005. Quo vadis ecosystem? Ecol. Model., 184(1): 5-17. https://doi.org/10.1016/j.ecolmodel.2004.11.005

Brown, J.H., K.V. Gupta, B.L. Li, B.T. Milne, C. Restrepo & G.B. West. 2002. The fractal nature of nature: power laws, ecological complexity and biodiversity. Philos. T. Roy. Soc. B., 357: 619-626. https://doi.org/10.1098/rstb.2001.0993

Brown, J.H., J.G. Gillooly, A.P. Allen, V.M. Savage & G.B. West. 2004. Towards a metabolic theory of ecology. Ecology, 85(7):1771-1789. https://doi.org/10.1890/03-0800

https://doi.org/10.1890/03-9000

Brown, J.H., A.P. Allen & J.F. Gillooly. 2007. The metabolic theory of ecology and the role of body size in marine and freshwater ecosystems, 1-10. In: Hildrew A.G., Raffaelli D.G. & R. Edmonds-Brown (Eds.) Body Size: the structure and function of aquatic ecosystems. Cambridge University Press, New York, 356 p. https://doi.org/10.1017/CBO9780511611223.002

Christensen, V. & D. Pauly, 1992. ECOPATH II - a software for balancing steady-state ecosystem models and calculating network characteristics. Ecol. Model., 61: 169-185 (www.ecopath.org) https://doi.org/10.1016/0304-3800(92)90016-8

Ernest, S.K.M., B.J. Enquist, J.H. Brown, E.L. Charnov, J.F. Gillooly, V.M. Savage, E.P. White, F.A. Smith, E.A. Hadly, J.P. Haskell, S.K. Lyons, B.A. Maurer, K.J. Niklas & B. Tiffney. 2003. Thermodynamic and metabolic effects on the scaling of production and population energy use. Ecol. Lett., 6: 990-995. https://doi.org/10.1046/j.1461-0248.2003.00526.x

Garlaschelli, D., G. Caldarelli & L. Pletonero. 2003. Universal scaling relations in food webs. Nature, 423: 165-168. https://doi.org/10.1038/nature01604

Jennings, S., K.J. Warr & S. Mackinson. 2002. Use of size-based production and stable isotope analyses to predict trophic transfer efficiencies and predator-prey body mass ratios in food webs. Mar. Ecol. Prog. Ser., 240, 11-20. https://doi.org/10.3354/meps240011

Jennings, S. & S. Mackinson. 2003. Abundancebody mass relationships in size-structured food webs. Ecol. Lett., 6(11): 971-974. https://doi.org/10.1046/j.1461-0248.2003.00529.x

Jennings, S., J.K. Pinnegar, N.V.C. Polunin & T.W. Boon. 2001. Weak cross-species relationships between body size and trophic level belie powerful size-based trophic structuring in fish communities. J. Anim. Ecol., 70(6): 934-944. https://doi.org/10.1046/j.0021-8790.2001.00552.x

Kitchell, J.F., T.E. Essington, C.H. Boggs, D.E. Schindler & C.J. Walters. 2002. The role of sharks and longline fisheries in a pelagic ecosystem of the Central Pacific. Ecosys� tems, 5(2): 202-216. https://doi.org/10.1007/s10021-001-0065-5

Lindeman, R.L. 1942. The trophic-dynamic aspect of ecology.Ecology, 23(4): 399-417. Available from: ftp://brd1.ucsc.edu/Bio293/ week2/Lindeman%201942.pdf [Accessed 7 June 2012] https://doi.org/10.2307/1930126

Merz, G. & R.A. Myers. 1998. A simplified formulation for fish production. Can. J. Fish.Aquat. Sci., 55(2): 478-484 https://doi.org/10.1139/f97-216

Pauly, D. & V. Christensen. 1993. Graphical representation of steady-state trophic ecosystem models, 20-28. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly (Eds.). Philippines, ICLARM Conf. Proc. 26. ISBN: 971-1022-84-2

Pauly, D. & V. Christensen. 1995. Primary production required to sustain global fisheries. Nature, 374: 255-257. https://doi.org/10.1038/374255a0

Savage, V.M., J.F. Gillooly, W.H. Woodruff, G.B. West, A.P. Allen, B.J. Enquist & J.H. Brown. 2004. The predominance of quarter-power scaling in biology. Funct. Ecol., 18(2): 257-282. https://doi.org/10.1111/j.0269-8463.2004.00856.x

Sokal, R.R. & F.J. Rolf. 1981. Biometry. 2nd ed., San Francisco, California, W.H. Freeman. ISBN: 0716712547

West, G.B. & J.H. Brown. 2004. Life's univer- sal scaling laws. Phys. Today, 57(9): 36-42.

https://doi.org/10.1063/1.1809090

West, G.B. & J.H. Brown. 2005. The origin of allometric scaling laws in biology from genomes to ecosystems: towards a quantitative unifying theory of biological structure and organization. J. Exp. Biol., 208: 1575-1592. https://doi.org/10.1242/jeb.01589

West, G.B., J.H. Brown & B.J. Enquist. 1997. A general model for the origin of allometric scaling laws in biology. Science, 276(5309): 122-126. https://doi.org/10.1126/science.276.5309.122

West, G.B., J.H. Brown & B.J. Enquist. 2001. A general model for ontogenic growth. Nature, 413: 628-631. https://doi.org/10.1038/35098076

(1) Okey, T.A. & D. Pauly, 1999. FCRR, 7(4): 137. Available from: http://www.fisheries. ubc.ca/publications/trophic-mass-balancemodel-alaskas-prince-william-sound-ecosystem-post-spill-period-199 [Accessed 1 February 2012]

(2) Dalsgaard, J. & D. Pauly, 1997. FCRR, 5(2): 33. Available from: http://www.fisheries. ubc.ca/publications/preliminary-mass-balance-model-prince-william-sound-alaskapre-spill-period-1980-1989 [Accessed 1 February 2012]

(3) Ainsworth, C., J.J. Heymans, T.J. Pitcher & M. Vasconcellos, 2002. FCRR, 10(4): 41. Available from: http://www.fisheries. ubc.ca/publications/ecosystem-models-northern-british-columbia-time-periods-2000-1950-1900-and-1750 [Accessed 1 February 2012]

(4) Morales-Zárate, M.V., F. Arreguín-Sánchez, J. López-Martínez & S.E. Lluch-Cota, 2004. Ecol. Model., 174: 331-345. https://doi.org/10.1016/j.ecolmodel.2003.09.028

(5) Arreguín-Sánchez, F., E. Arcos & E.A. Chávez, 2002. Ecol. Model., 156: 167-183. https://doi.org/10.1016/S0304-3800(02)00159-X

(6) Salcido-Guevara, L.A., 2006. M.Sc. thesis, CICIMAR-IPN., 83 Available from: http:// www.biblioteca.cicimar.ipn.mx/oacis/tesisdesplegardetalles.php?id=422 [Accessed 1 February 2012]

(7) Wolff, M., V. Koch, J.B. Chavarria & J. Vargas, 1998. Available from: http://www. cimar.ucr.ac.cr/volumenes-especiales/contenidos/cont_eacr1.html [Accessed 1 February 2012]

(8) Wolf, M., H.J. Hartmann & V. Koch, 1996. Rev. Biol. Trop., 44(3): 215-231. Available from: http://www.cimar.ucr.ac.cr/volumenes-especiales/contenidos/cont_hensen. html [Accessed 2 February 2012]

(9) Olivieri, R.A., A. Cohen & F.P. Chavez, 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 315-322. ISBN: 9711022842

(10) Arreguín-Sánchez, F., A. Hernández-Herrera, M. Ramírez-Rodríguez & H. PérezEspaña, 2004. Ecol. Model., 172: 373-382.

(11) Watkinson, S. & D. Pauly, 1999. Changes in the ecosystem of Rivers Inlet, British Columbia: 1950 vs. the present. Available from: http://www.ecopath.org/model/51 [Accessed 1 February 2012] 12)

(12) Zetina-Rejón, M.J., F. Arreguín-Sánchez & E.A. Chávez, 2003. Estuar.Coast. Shelf Sci., 57: 803-815. https://doi.org/10.1016/S0272-7714(02)00410-9

Kitchell, J.F., T.E. Essington, C.H. Boggs, D.E. Schindler & C.J. Walters. 2002. Eco� systems, 5(2): 202-216. https://doi.org/10.1007/s10021-001-0065-5

(14) Zetina-Rejón, M.J. & F. Arreguín-Sánchez, 2002. In: Wakida-Kusunoki, A., R. SolanaSansores & J. Uribe-Martínez, (Eds.) Mexico, INP, 49-56. Available from: http://www. inapesca.gob.mx/portal/documentos/publ icaciones/11Tercer+foro+de+camaron.pdf [Accessed 2 February 2012]

(15) Morissette, L., D. Simon-Pierre, C. Savenkoff, M.O. Hammill, H. Bourdages &D. Chabot, 2003. Can. Tech. Rep. Fish.Aqua. Sci., 2497: 94. Avalaible from: http://www. dfo-mpo.gc.ca/Library/276401.pdf [Accessed 2 February 2012]

(16) Christian, R.R. & J.J. Luczkovich, 1999. Ecol. Model., 117(1): 99-124. https://doi.org/10.1016/S0304-3800(99)00022-8

(17) Vidal, L. & M. Basurto, 2003. FCRR, 11(6): 255-264. Available from: http://www. fisheries.ubc.ca/publications/mexico-brazilcentral-atlantic-fisheries-catch-trends-andecosystem-models [Accessed 2 February 2012]

Vega-Cendejas, M.E. & F. Arreguín-Sánchez, 2001. Ecol. Model., 137: 119-133. https://doi.org/10.1016/S0304-3800(00)00421-X

(19) E. Rivera-Arriaga, A. L. Lara-Domínguez, G. Villalobos-Zapata & A. Yáñez-Arancibia, 2003. FCRR, 11(6): 245-254. Available from: http://www.fisheries.ubc.ca/publications/mexico-brazil-central-atlantic-fisheries-catch-trends-and-ecosystem-models [Accessed 2 February 2012]

(20) Venier, J.M. & D. Pauly, 1997. Proc. 8th Int. Coral Reef Sym., 1: 915-920. Available from: http://www.ecopath.org/model/64 [Accessed 2 February 2012]

(21) Alvarez-Hernández, J.H., 2003. FCRR, 11(6): 227-235. Available from: http://www. fisheries.ubc.ca/publications/mexico-brazil central-atlantic-fisheries-catch-trends-andecosystem-models [Accessed 2 February 2012]

(22) Halfon, E. & N. Shito, 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 29-39. ISBN: 9711022842

(23) Duarte, L.O. & C.B. García, 2004. Ecol. Model., 172: 323-338. https://doi.org/10.1016/j.ecolmodel.2003.09.014

(24) Pedersen, S.A, 1994. ICES, 26. Available from: http://www.ecopath.org/model/48 [Accessed 2 February 2012]

(25) Manickchand-Heileman, S., L.A. Soto & E. Escobar, 1998. Estuar. Coast. Shelf Sci., 46(6): 885-899. https://doi.org/10.1006/ecss.1997.0324

(26) Manickchand-Heileman, S., F. ArreguínSánchez, A. Lara-Dominguez & L.A. Soto, 1998. J. Fish Biol., 53: 179-197. (1998). https://doi.org/10.1111/j.1095-8649.1998.tb01026.x

(27) De La Cruz-Agüero, G., 1993. In: Tro� phic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 193-196. ISBN: 9711022842

(28) Rosado-Solórzano, R. & S.A. Guzmán del Próo, 1998. Ecol. Model., 109(2): 141-154. https://doi.org/10.1016/S0304-3800(98)00011-8

(29) Chávez, E.A., M. Garduño & F. ArreguínSánchez, 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 186-192. ISBN: 9711022842

(30) Buchary, E.A., 2001. FCRR, 9(4): 350: 198-206. Available from: http://www.fisheries.ubc.ca/publications/fisheries-impactsnorth-atlantic-ecosystems-models-andanalyses [Accessed 2 February 2012]

(31) Rybarczyk, H. & B. Elkaïm, 2003. Estuar. Coast. Shelf Sci., 58(4): 775-791. https://doi.org/10.1016/S0272-7714(03)00184-7

(32) Carrer, S.& S. Opitz. Ecol. Model, 1999. 124: 193-219. https://doi.org/10.1016/S0304-3800(99)00161-1

(33) Lobry, J. 2004. Thesis doctoral, France, Université de Bordeaux I. Available from: http://ori-oai.u-bordeaux1.fr/ori-oaisearch/notice.html?id=u-bordeaux1-ori- 901&format=dc_id [Accessed 2 February 2012]

(34) Palomares, M.L., P. Reyes-Marchant, N. Lair, M. Zainure, G. Barnabe & G. Lasserre, 1993. In: Trophic models of aquatic eco� systems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 224- 229. ISBN: 9711022842

(35) Brando, V.E., R. Ceccarelli, S. Libralato & G. Ravagnan. 2004. Ecol. Model., 172: 213-232. https://doi.org/10.1016/j.ecolmodel.2003.09.008

(36) Palomares, M.L., B. Yulianto, L. Puy, D. Bengen & A. Belaud, 1993. In: Trophic mod� els of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 172-179. ISBN: 9711022842

(37) Reyes-Marchant, P., J.L. Jamet, N. Lair, H. Taleb & M.L.D. Palomares, 1993. In: Tro� phic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 95-101. ISBN: 9711022842

(38) Gamito, S. & K. Erzini, 2005. Ecol. Model., 181(4): 509-520. (2005). https://doi.org/10.1016/j.ecolmodel.2004.02.024

(39) Harvey, C.J., S.P. Cox, T.E. Essington, S. Hansson & J.F. Kitchell, 2003. ICES J. Mar. S., 60: 939-950. https://doi.org/10.1016/S1054-3139(03)00098-5

(40) Walline, P.D., S. Pisanty, M. Gophen & T. Berman, 1993. In: Trophic models of aquat� ic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 103-109. ISBN: 9711022842

(41) Kolding, J., 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 116-123. ISBN: 9711022842

(42) Moreau, J., V. Christensen & D. Pauly, 1993. In: Trophic models of aquatic eco� systems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 124- 129. ISBN: 9711022842

(43) Moreau, J., W. Ligtvoet & M.L.D. Palomares, 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 144-152. ISBN: 9711022842

(44) Moreau, J., B. Nyakageni, M. Pearce & P. Petit, 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 138-143. ISBN: 9711022842

(45) Machena, C., J. Kolding & R.A. Sanyanga 1993. In: Trophic models of aquatic eco� systems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 130- 137. ISBN: 9711022842

(46) Degnbol, P., 1993 In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 110-115. ISBN: 9711022842

(47) Moreau, J., M.C. Villanueva, U.S. Amarasinghe & F. Schiemer, 2001. In: Reservoir and culture-based fisheries: biology and management, De Silva, S.S., (Ed.). Australia, Aust. Centre Inter. Agr. Res., 201-214. ISBN: 0642456941

(48) Silvestre, G., S. Selvanathan & A.H.M. Salleh, 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 300-306. ISBN: 9711022842

(49) Garces, L.R., M. Alias, A. A. Talib, M. Mohamad-Norizam & G. T. Silvestre, 2003. In: Silvestre, G., L. Garces, I. Stobutzki, C. Luna, M. Ahmed, R.A.V. Santos, L. Lachica-Aliño, P. Munro, V. Christensen &D. Pauly, (Eds.). Assessment, Management and Future Directions for Coastal Fisheries in Asian Countries.WorldFish Center. 333- 352. ISBN: 9832346223

(50) Hsing-Juh, L., S. Kwang-Tsao, H. JiangShiou, L. Wen-Tseng, C. I-Jiunn & L. LihHuwa, 2004. J. Mar. Sci. Tech., 12(5): 424- 432. Available from: http://fishdb.sinica. edu.tw/pdf/116.pdf [Accessed 2 February 2012]

(51) Bundy, A. & D. Pauly, 2001. Fish. Res., 53(3): 263-281. https://doi.org/10.1016/S0165-7836(00)00295-2

(52) W.L. Campos, 2003. In: Silvestre, G., L. Garces, I. Stobutzki, C. Luna, M. Ahmed, R.A.V. Santos, L. Lachica-Aliño, P. Munro, V. Christensen & D. Pauly, (Eds.). Assess� ment, Management and Future Direc� tions for Coastal Fisheries in Asian Coun� tries.WorldFish Center. 353-364. ISBN:9832346223

(53) Aliño, P.M., L.T. McManus, J.W. Nañola, J.M.D. Fortes, G.C. Trono & G.S. Jacinto, 1993. In: Trophic models of aquatic eco� systems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 252-258. ISBN: 9711022842

(54) Delos Reyes, M.R., 1995. Thesis doctoral, Germany, Universität Hamburg, 138. Available from: https://kataloge.uni-hamburg.de/ DB=1/SET=4/TTL=1/LNG=EN/NXT [Accessed 2 February 2012]

(55) Man, A., 2003. In: Silvestre, G., L. Garces, I. Stobutzki, C. Luna, M. Ahmed, R.A.V. Santos, L. Lachica-Aliño, P. Munro, V. Christensen & D. Pauly, (Eds.). Assess� ment, Management and Future Direc� tions for Coastal Fisheries in Asian Coun� tries.WorldFish Center. 313-332. ISBN: 9832346223

(56) Nurhakim, S., 2003. In: Silvestre, G., L. Garces, I. Stobutzki, C. Luna, M. Ahmed, R.A.V. Santos, L. Lachica-Aliño, P. Munro, V. Christensen & D. Pauly, (Eds.). Assess� ment, Management and Future Direc� tions for Coastal Fisheries in Asian Coun� tries.WorldFish Center. 299-312. ISBN:9832346223

(57) Tong, L., Q. Tang & D. Pauly, 2000. Chi� nese J. App. Ecol., 11(3): 435-440. Available from: http://www.cjae.net/EN/volumn/ volumn_1315.shtml [Accessed 2 February 2012]

(58) Golam-Mostafa, M., 2003. In: Silvestre, G., L. Garces, I. Stobutzki, C. Luna, M. Ahmed, R.A.V. Santos, L. Lachica-Aliño, P. Munro, V. Christensen & D. Pauly, (Eds.). Assess� ment, Management and Future Direc� tions for Coastal Fisheries in Asian Coun� tries.WorldFish Center. 263-280. ISBN: 9832346223

(59) Gribble, N.A., 2003. Fish. Res., 65: 493- 506. https://doi.org/10.1016/j.fishres.2003.09.035

(60) Tudman, P.D., 2001. Thesis, Australia, James Cook University. Available from: http://www.ecopath.org/model/77 [Accessed 2 February 2012]

(61) Bozec, Y.M. D. Gascuel & M. Kulbicki, 2004. Aqua. Living Resour., 17(2): 151- 162. https://doi.org/10.1051/alr:2004024

(62) Cox, S.P., T.E. Essington, J.F. Kitchell, S.J.D. Martell, C.J. Walters, C. Boggs & I. Kaplan, 2002. Can. J. Fish.Aquat. Sci., 59: 1736-1747. https://doi.org/10.1139/f02-138

(63) Bradford-Grieve, J.M., P.K. Probert, S.D. Nodder, D. Thompson, J. Hall, S. Hanchet, P. Boyd, J. Zeldis, A.N. Baker, H.A. Best, N. Broekhuizen, S. Childerhouse, M. Clark, M. Hadfield, K. Safi & I. Wilkinson, 2003. J. Exp. Mar. Biol. Ecol., 289(2): 223-262. https://doi.org/10.1016/S0022-0981(03)00045-5

(64) Neira, S., H. Arancibia & L. Cubillos, 2004. Ecol. Model., 172: 233-248. https://doi.org/10.1016/j.ecolmodel.2003.09.009

(65) Mendy, A.N., 1999. Reykjavik, United Nations University. Available from: http://www. ecopath.org/model/58 [Accessed 2 February 2012]

(66) Lee Bredesen, E., 1999. Thesis, Canada, University of British Columbia. Available from: http://www.ecopath.org/model/70 [Accessed 2 February 2012]

(67) Heymans, J.J. & D. Baird, 2000. Ecol. Model., 131: 97-119. https://doi.org/10.1016/S0304-3800(00)00275-1

(68) Aydin, K.Y., V.V. Lapko, V.I. Radchenko & P.A. Livingston, 2002. NOAA Technical Memorandum. Available from: http://www. afsc.noaa.gov/Publications/techmemos_ older.htm [Accessed 2 February 2012]

(69) Mendoza, J. J., 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 285-297. ISBN: 9711022842

(70) Vasconcellos, M. & M.A. Gasalla, 2001. Fish. Res., 50(3): 279-295. https://doi.org/10.1016/S0165-7836(00)00217-4

(71) Blanchard, J.L., J.K. Pinnegar & S. Mackinson, 2002. UK, CEFAS, Sci. Ser. Tech. Rep., Available from: http://www.cefas. defra.gov.uk/publications/scientific-series/ technical-reports.aspx [Accessed 2 February 2012]

(72) Vivekanandan, E., M. Srinath, V.N. Pillai, S. Immanuel & K.N. Kurup, 2003. In: Silvestre, G., L. Garces, I. Stobutzki, C. Luna, M. Ahmed, R.A.V. Santos, L. Lachica-Aliño, P. Munro, V. Christensen & D. Pauly, (Eds.). Assessment, Management and Future Di� rections for Coastal Fisheries in Asian Countries.WorldFish Center. 281-298. ISBN: 9832346223

(73) Ortiz, M. & M. Wolf. Aquatic Conservation: Marine and Freshwater Ecosystems. 12(6): 601-618. https://doi.org/10.1002/aqc.512

(74) Ryzbarczyk, H., B. Elkaim, L. Ochs & N. Loquet, 2003. Estuar. Coast. Shelf Sci., 58(3): 405-421. https://doi.org/10.1016/S0272-7714(02)00294-9

(75) De Paula E Silva, R. & A.M. Caramelo., 1993. In: Trophic models of aquatic eco� systems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 214- 223. ISBN: 9711022842

(76) Pauly, D., 2002. In: The Gulf of Guinea Large Marine Ecosystem, McGlade, J.M., P. Cury, K.A. Koranteng, &N.J. HardmanMountford, (Eds.). Elsevier, 289-295. ISBN: 0444510281

Vega-Cendejas, M.E., 2003. FCRR, 11(6): 237-243. Available from: http://www.fisheries.ubc.ca/publications/mexico-brazilcentral-atlantic-fisheries-catch-trends-andecosystem-models [Accessed 2 February 2012]

(78) Lin, H.J., K.T. Shao, S.R. Kuo, H.L. Hsieh, S.L. Wong, I.M. Chen, W.T. Lo & J.J. Hung, 1999. Estuar.Coast. Shelf Sci., 48(5): 575- 588. https://doi.org/10.1006/ecss.1998.0457

(79) Gribble, N.A., 2001. In: Oceanographic Processes of Coral Reefs: Physical and Biological Links in the Great Barrier Reef, Wolanski, E., (Ed.). CRC Press, 189-207. ISBN: 084930833X. https://doi.org/10.1201/9781420041675.ch12

(80) Arias-González, J.E., B. Delesalle, B. Salvat & R. Galzin, 1997. Coral Reefs, 16(4):231-246. https://doi.org/10.1007/s003380050079

(81) Heymans, S.J.J. 2005. FCRR, 13: 8-82. Available from: http://www.fisheries.ubc.ca/ publications/food-web-models-and-datastudying-fisheries-and-environmental-impacts-eastern-pacific-e [Accessed 2 February 2012]

(82) Mathews, C.P., 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 161-171. ISBN: 9711022842

(83) Angelini, R. & A.A. Agostinho, 2005. Ecol. Model., 181: 109-121. https://doi.org/10.1016/j.ecolmodel.2004.06.025

(84) Aravindan, C.M., 1993. In: Trophic models of aquatic ecosystems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 87-89. ISBN: 9711022842

(85) Palomares, M.L.D., K. Horton & J. Moreau, 1993. In: Trophic models of aquatic ecosys� tems, Christensen, V. & D. Pauly, (Eds). Philipines, ICLARM Conf. Proc. 26., 153- 158. ISBN: 9711022842

Descargas

Publicado

2012-12-04

Cómo citar

Salcido -Guevara, L. A., Arreguí­n -Sánchez, F., Palmeri, L., & Barausse, A. (2012). Regularidad del escalamiento metabólico en ecosistemas acuáticos. CICIMAR Oceánides, 27(2), 13–26. https://doi.org/10.37543/oceanides.v27i2.113

Número

Sección

Artículos